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Volume 46, Number 1, February 2000:
The Permian Insect Fossils of Elmo, Kansas

Image - front cover of issue

ISSUE HOME PAGE

ABOUT THIS ISSUE
- about KSN
- about the author

IN THIS ISSUE
- introduction
- the elmo site as part of the Kansas-Oklahoma permian
- a short history of the discovery and study of the elmo fossils
- the insects: part 1, part 2, part 3,
- references
- back cover

SLIDESHOW
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This page was last modified:
October 15, 2003

Originally posted:
March 20, 2003


 

The Permian Insect Fossils of Elmo, Kansas
by Roy J. Beckemeyer


The Insects (cont'd)


Uncertain Order (Pterygota: Palaeoptera) 
One genus, 1 species.
- Uncertain Family
-- Permoneura Carpenter 1931
---- lameerei Carpenter 1931

Protorthoptera (Pterygota: Neoptera: Exoptery-gota) U.Carb.-Triass.  This is a large group of orthopteroid insects that dominated the entomofauna of the Paleozoic and is the most common order found at Elmo.  Many of the taxa (especially the family Lemmatophoridae) are obviously related to today’s stoneflies (Order Plecoptera) and were probably aquatic in their immature stages.  There are many species for which even body parts were well preserved. Thirteen families, 24 genera, 34 species.
-
Blattinopsidae Bolton 1925
-- Blattinopsis Giebel 1867
---- kukalovae Carpenter 1966
-- Glaphyrophlebia Handlirsch 1906
---- speciosa (Sellards) 1909 (Sindon)
---- ovata (Sellards) 1909 (Pursa)
- Chelopteridae Carpenter 1950
-- Chelopterum Carpenter 1950
-- peregrinum Carpenter 1950


Figure 13 - L. typa after Carpenter 1935 (Ref. 10)
Figure 13.  L. typa  after Carpenter 1935 (Ref. 10)
Figure 14 - L. typa fossil

Fig. 14.  L. typa fossil from teaching collection of Prof. George Byers of the University of Kansas.  Fossil identified and image scanned by Roy Beckemeyer, Oct. 1999.

- Demopteridae Carpenter 1950
--
Demopterum Carpenter 1950
---- gracile Carpenter 1950
- Heteroptilidae Carpenter 1976
-- Heteroptilon Carpenter 1976
---- costale Carpenter 1976
- Lemmatophoridae Sellards 1909
-- Lemmatophora Sellards 1909
---- typa Sellards 1909 (Fig’s. 13, 14)
-
Artinska Sellards 1909
---- clara Sellards 1909
---- sellardsi Tillyard 1928
---- ovata  (Sellards) 1909 (Orta)
-- Lisca Sellards 1909
---- minuta Sellards 1909
-- Paraprisca Handlirsch 1919
---- fragilis (Sellards) 1909 (Prisca)
---- grandis Carpenter 1935
-- Lecorium Sellards 1909
---- elongatum Sellards 1909
- Liomopteridae Sellards 1909
-- Liomopterum Sellards 1909
---- ornatum Sellards 1909
----- elongatum (Sellards) 1909 (Horates)
----- sellardsi Carpenter 1950
-- Semopterum Carpenter 1950
---- venosum Carpenter 1950
-- Tapopterum Carpenter 1950
---- celsum Carpenter 1950
- Phenopteridae Carpenter 1950
-- Phenopterum Carpenter 1950
---- elongatum (Sellards) 1909 (Lepium)
- Probnidae Sellards 1909
-- Probnis Sellards 1909
---- speciosa Tillyard 1939

 

Figure 15 - P. permiana after Carpenter 1950 (Ref. 14)

Fig. 15.  P. permiana after Carpenter 1950 (Ref. 14)

 
- Protembiidae Tillyard 1937
-- Protembia Tillyard 1937
----- permiana Tillyard 1937 (Fig. 15)
-
Psoropteridae Carpenter 1976
-- Psoroptera Carpenter 1976
---- cubitalia Carpenter 1976
- Steropteridae Carpenter 1950
-- Steropterum Carpenter 1950
---- rotundum Carpenter 1950
---- maculosum Carpenter 1966
----- breve Carpenter 1966
- Strephocladidae Martynov 1938
-- Homocladus Carpenter 1966
---- grandis Carpenter 1966
---- ornatus Carpenter 1966
-- Paracladus Carpenter 1966
---- retardatus Carpenter 1966
- Tococladidae Carpenter 1966
-- Tococladus Carpenter 1966
---- rallus Carpenter 1966
-- Opisthocladius Carpenter 1976
---- arcuatus Carpenter 1976
----- strictus Carpenter 1976
- Uncertain Family
-- Atava Sellards 1909
---- ovata Sellards 1909
-- Lecopterum Sellards 1909
---- delicosum Sellards 1909

Blattaria (Pterygota: Neoptera: Exopterygota) U.Carb.-Hol.  The cockroaches were one of the largest orders of insects in the late Paleozoic.  Carpenter (Ref. 19) did not revise the nomenclature for this order in his Treatise, so the taxonomy is listed here as defined by Tillyard.  One family, 2 genera, 4 species.

- Spiloblattinidae Handlirsch
-- Permoblattina Tillyard 1937
---- curta (Sellards) 1908 (Etoblattina)
---- permiana (Sellards) 1908 (Etoblattina)
-- Pycnoblattina Sellards 1908
---- compacta (Sellards) 1908 (Puknoblattina)
---- curvata (Sellards) 1908 (Puknoblattina)

Protelytroptera (Pterygota: Neoptera: Exoptery-gota) Perm.-Cret.  An order of small insects that are related to cockroaches and are ancestral to modern earwigs (Order Dermaptera).  The front wings, called elytra, are actually hard covers like those of beetles.  Five families, 9 genera, 15 species.

- Archelytridae Carpenter 1933
-- Archelytron Carpenter 1933
---- superbum Carpenter 1933
- Elytroneuridae Carpenter 1933
-- Elytroneura Carpenter 1933
---- permiana Carpenter 1933
- Megelytridae Carpenter 1933
-- Megelytron Tillyard 1931
---- robustum Tillyard 1931
- Permelytridae Tillayrd 1931
-- Permelytron Tillyard 1931
---- schucherti  Tillyard 1931
-- Blattelytron Tillyard 1931
---- permianum Tillyard 1931
-- Parablattelytron Tillyard 1931
---- subincisum Tillyard 1931
---- rectum Tillyard 1937
---- elongatum (Tillyard) 1931 (Acosmelytron)
---- delicatum (Tillyard) 1931 (Acosmelytron)
---- latum Carpenter 1937

Figure 16 - Protelytron permianum after Carpenter & Kukalova, 1964 (Ref. 21)

Fig. 16.  Protelytron permianum after Carpenter & Kukalova, 1964 (Ref. 21)

  
- Protelytridae Tillyard 1931
-- Protelytron Tillyard 1931
---- permianum Tillyard 1931 (Fig. 16)
---- angustum Carpenter 1933
---- furcatum Carpenter 1939
-- Permelytropsis Carpenter 1933
---- cubitalis  Carpenter 1933
- Uncertain Family
-- Protelytropsis Tillayrd 1931
---- grandis Tillyard 1931

Orthoptera (Pterygota: Neoptera: Exopterygota) U.Carb.-Hol.  Carpenter placed in this order only the fossils of saltatorial, or leaping, insects similar to our grasshoppers.  Four families, 5 genera, 7 species.

- Oedischiidae Handlirsch 1906
-- Paroedischia Carpenter 1966
---- rectu Carpenter 1966
---- maculata Carpenter 1966
- Permelcanidae Sharov 1962
-- Promartynovia Tillyard 1937
---- venicosta Tillyard 1937
- Anelcanidae Carpenter 1987
-- Anelcana Carpenter 1987
---- dilatata (Carpenter) 1966 (Parelcana)
-- Petrelcana Carpenter 1966
---- elongata Carpenter 1966
- Permoraphididae Tillyard 1932
-- Permoraphidia Tillyard 1932
---- americana Tillyard 1932
---- grandis Carpenter 1943

Psocoptera (Pterygota: Neoptera: Exopterygota) Perm.-Hol.  Psocids are still found today and are called bark lice or book lice.  They are diminutive insects with large heads, prominent eyes, and chewing mouthparts.  Three families, 7 genera, 12 species.

- Psocidiidae Tillyard 1926
-- Dichentotum Tillyard 1926
---- tinctum Tillyard 1926  (Fig. 17)
---- minimum (Tillyard) 1926
---- latum Carpenter 1932
---- parvulum Carpenter 1932
---- grande Carpenter 1933
- Permopsocidae Tillyard 1926
-- Prognopsocus Tillyard 1926
---- permianus Tillyard 1926
-- Permopsocus Tillyard 1926
---- latipennis Tillyard 1926
---- ovatus Carpenter 1939
-- Lithopsocidium Carpenter 1932

Fig. 17.  D. tinctum after Carpenter 1933 (Ref. 9)

 



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